Mary Jean “Lily” Tomlin (born September 1, 1939) is an American actor, comedian, writer and producer. During her 40-year career she has also been nominated for an Academy Award, and has won mutiple awards from many quarters, including Tony Awards, Emmy Awards, and a Grammy Award.

In 1969, Tomlin joined the sketch comedy show Laugh-In. Some characters from the show have been associated with her throughout her career, including the gum-chewing, wisecracking, snorting telephone operator, Ernestine; the bratty five-year-old Edith Ann, seated in an over-sized rocking chair making rude noises while telling stories about her baby brother and pet dog Buster; and the Tasteful Lady, who lives a gracious, nave life of entitlement in the upper class and shades of whom show up in Tomlin's film role in All of Me. Additional characters include Susie the Sorority Girl, who appeared on Tomlin's album Modern Scream and in her 1975 appearance on Saturday Night Live.

Tomlin was also one of the first female comedians to break out in male drag. Though drag had been around in Hollywood for some time by men, Tomlin broke new ground by not only crossing gender stereotypes, but racial ones as well. She accomplished this in the late 70's with Pervis Hawkins, a black rhythm-and-blues soul singer (patterned after Luther Vandross), with a mustache, beard and close-cropped afro hairstyle, dressed in a three-piece suit. Tomlin used very little if any skin-darkening cosmetics (it usually depended on stage lighting) as part of the character.

AT&T offered Tomlin $500,000 to play her character Ernestine in a commercial, but she declined saying it would compromise her artistic integrity. However, in 1976 she did appear as Ernestine in a parody of a commercial on Saturday Night Live , in which she proclaimed, "We don't care, we don't have to...we're the phone company." The character would later make a guest appearance at The Superhighway Summit at UCLA, January 11, 1994, interrupting a speech being given on the information superhighway by then-Vice President Al Gore. In 2003, she made two commercials as Ernestine for WebEx.

Tomlin is noted for her versatility. In Robert Altman's Nashville, for which she was nominated for an Academy Award for Best Supporting Actress, she played Linnea Reese, a straitlaced, gospel-singing, mother of two deaf children who has an affair with a country singer (played by Keith Carradine). The Oscar that year went to Lee Grant for her role in Shampoo. She was also a secretary Violet Newstead in Nine to Five, performed several comedic roles in the 1981 film The Incredible Shrinking Woman, and was a sickly heiress in the Steve Martin comedy All of Me.

She and Bette Midler played two pairs of identical twins who were switched at birth in the 1989 comedy Big Business. Tomlin also played chain-smoking waitress Doreen Piggott in Altman's 1993 ensemble film Short Cuts, and, in two films by director David O. Russell; she appeared as a peacenik Raku artist in Flirting with Disaster and later, as an existential detective in I ♥ Huckabees. In 2007, a video recording surfaced showing Tomlin and Russell in a heated exchange over the shooting of a scene in Huckabees.

Tomlin voiced Ms. Frizzle on the animated television series The Magic School Bus from 1994 to 1998. Also, in the 1990s, Tomlin appeared on the popular sitcom Murphy Brown as the title character's boss. In 2005 and 2006, she had a recurring role as Will Truman's boss Margot on Will & Grace. She appeared on the dramatic series The West Wing for four years (2002-2006) in the recurring role of presidential secretary Deborah Fiderer.

Tomlin starred in the 1985 hit one-woman Broadway show The Search For Signs of Intelligent Life in the Universe, written by her long-time life partner, writer/producer Jane Wagner. The show won her a Tony Award, and was made into a feature film in 1991. Tomlin revived the show for a brief run in 2000. In 1989, she won the Sarah Siddons Award for her work in Chicago theatre.

She collaborated again with director Robert Altman, starring in the film A Prairie Home Companion, playing half of a middle-aged Midwestern singing duo with Meryl Streep.

In the 2008-2009 fifth season of Desperate Housewives she has a recurring role as Roberta, the sister of Mrs. McCluskey (played by Kathryn Joosten). Previously on The West Wing, Tomlin had played the successor to Delores Landingham, the previous secretary, who was played by Joosten.

During the 2008 Emmy Awards, Tomlin appeared as part of a tribute to the seminal 1960s television series Laugh-In. Tomlin voiced Tammy in the 2005 The Simpsons episode, "The Last of the Red Hat Mamas." Tomlin will provide a voice for the upcoming film Ponyo on the Cliff by the Sea, projected for an April 2009 release, although the specific role is as yet unknown.